By Hans A. Krebs, H.L. Kornberg, K. Burton
This survey used to be written on the invitation of the Editors of the "Ergebnisse der Physiologie". Its objective is to provide the more moderen development within the understand ledge of organic strength differences. because it used to be meant for a assessment magazine, the reader used to be taken to be accustomed to the basics of present biochemistry, as defined within the common textbooks. It used to be now not the thing to bring together an intensive selection of proof. The survey is proscribed to features of wider curiosity, and the most emphasis has been at the normal unifying rules which emerge from the good mass of specific ob servations. the object is reprinted within the wish that it can be precious during this shape to complex scholars and examine employees in biochemistry and similar matters. H. A. KREBS H. L. KORNBERG 2 desk of Contents web page 1. the foremost place of Adenosine Triphosphate . . . 213 2. the 3 stages of food Degradation. . . . 213 three. The Energy-Yielding Steps of middleman Metabolism 215 four. The Build-up of Phosphate Bond strength ..... 221 five. replacement Pathways of Anaerobic Fermentation in Micro-organisms. 227 6. replacement Pathways of Glucose Oxidation . 237 7. the trail of Carbon in Photosynthesis . . . 243 eight. usage of power for Chemical Syntheses 249 nine. keep an eye on of Energy-Supplying tactics . . . 262 10. a different function of ATP as an strength shop. 271 eleven. Evolution of power remodeling Mechanisms 273 Appendix via okay. BURTON unfastened power info of organic curiosity 27S References . . . . . . . . . . . . . . .
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Additional resources for Energy Transformations in Living Matter: A Survey
In the glucose breakdown by fermentation followed by the tricarboxylic acid cycle there are at least 6 different reactions leading to the reduction of pyridine nucleotide, against 2 in the pentose phosphate cycle. The total number of steps by which the complete oxidation of glucose is achieved is smaller in the pentose phosphate cycle than in the breakdown by fermentation plus tricarboxylic acid cycle, but the latter route not only covers the degradation of carboJC02 Jco2 hydrate but also that of fats and amino acids.
1··· I O=C-H H--1-oH + HOC-H ...... OPO*H. ~H,OPOIH. H. H. plwspAale The fructose 6-phosphate fonned in (6, 6) and (6, 7) is converted to glucose 6-phosphate by reaction (6,8), catalysed by hexosephosphate isomerase. H, glmose 6-Phosphate This reaction completes the cycle in that it leads to the (partial) regeneration of the starting material, glucose 6-phosphate. The interplay of the components of the cycle is somewhat complex. It is shown diagrammatically in Fig. 7. _. P Fig. 1. DiAgram 01 lite pmlose phosp1late eye/4 = Starting materials and end products.
The sedoheptulose 1: 7-diphosphate produced is next (7, 5) postulated to yield sedoheptulose 7-phosphate, by hydrolysis of the phosphate group attached to carbon atom 1 CH 20H CHP POaH 2 I I c=o I HO-C-H I H-C-OH I H-C-OH I H-C-OH I CHP POaH 2 sedoheptulose 1: 7-diphosphate C=O I I H-C-OH I H-C-OH I H-C-OH I HO-C-H CHPP03:ti2 sedoheptulose 7-phosphate (7, 5) 248 H. A. KREBS and H. L. KORNBERG: Energy Transformations in Living Matter under the action of a phosphatase. Again, this reaction is similar to the conversion of fructose 1: 6-diphosphate to fructose 6-phosphate already described.